True, but, if I recall correctly, many cells use the electrical potential difference between the inside and outside of the cell as the driving force for many active transport systems, or would that be passive transport since no ATP is required?
Excess negative charges outside the cell would alter this potential, changing the effectiveness of the transport mechanism. Just a shot in the dark here, btw. If the cells were less effective, or had to resort to energy requiring pumps, then perhaps the extra energy required for cell homeostasis would result in less energy devoted to growth.
This could also be the case in cells regulating their internal PO4 content, with a steeper up-concentration gradient to work against.
But again, PO4 could simply be a measurable proxy for other organics more prevalent in overstocked tanks that could cause unhealthy conditions, reducing growth rates...
Molecular Biology aside, it seems reasonable that the Ca++ & PO4-- interactions would increase with PO4 concs. With Ca more difficult to obtain for bone development, nerve and muscle control, this too could reduce growth, I think.
Man it's been a long time since my last bio class...