Hmm......so what does it mean when the agressor does blind-side hits and then follows them up with lip-locking? Oh, and then throws in charges with fins flared and head shaking? That's the behavior I saw between my two angels. BTW, the loser didn't just cower at first. He/she did fight back. For a brief time, it was the loser that was doing the chasing and attacking.......I mean for about a day or so. For whatever reason, the dominant player was a complete wuss for that day.
You have a good chance of a fish deciding territory..most likely a male(not always)
keep a close eye out..angels can hit pretty hard ..I have seen some hit's so hard they stun the lesser fish.
I have also lot angels from this behavior.