As the organic detritus sinks into the DSB and is eaten by bacteria, oxygen is consumed. The more detritus, the more O2.
That's what I thought was causing my fish to gasp, till you told me about CO2 excess. I was glad to hear that, because low oxygen would have been an ironic problem for a girl whose forum name means, "no oxygen", LOL.
I found a link to the "anoxic filtration" method gaining popularity in koi keeping, it uses clay, but it is in baked form (kitty litter) so you don't get the turbidity problem that regular clay can cause.
http://www.essexsection.co.uk/html/anoxic_filtration_system.html